Keith’s strengths are in the areas of freshwater ecology and aquatic ecotoxicology, with an emphasis in quantitative methods.  He is a broadly trained freshwater ecologist, with 20 years of field and laboratory experience in algal, invertebrate, and fish ecology; general community ecology; and freshwater biogeochemistry. Since joining GLEC in 2009, he has supported development of EPA water quality criteria documents for selenium, ammonia, cadmium, chloride, aluminum, and triclosan; and he has authored or coauthored numerous additional technical support documents, white papers, and technical reports.  Keith is proficient with several statistical packages, including SYSTAT and R.  His doctoral research included investigations into the effects of sediment toxicity, particularly metal bioavailability; the effects of watershed management practices on stream ecosystems; the effects of chemical toxicants on stream ecosystems; and the interactive effects of light, nitrogen, and phosphorus on stream ecosystems.  Keith holds a M.S. in Ecology from the University of North Carolina and a Ph. D. in Ecology, Evolution, and Marine Biology form the University of California, Santa Barbara.